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BZN Volume 58, Part 3, 28 September 2001

General Articles & Nomenclatural Notes


General Articles and Nomenclatural Notes with the following titles were published on 28 September 2001 in Volume 58, Part 3 of the Bulletin of Zoological Nomenclature

Copies of these General articles and Nomenclatural Notes can be obtained free of charge from the Executive Secretary, The International Commission on Zoological Nomenclature, c/o The Natural History Museum, Cromwell Road, London SW7 5BD, U.K. (e-mail: iczn@nhm.ac.uk).


General Article

Zoological nomenclature - reflections on the recent past and ideas for our future agenda

Alessandro Minelli
Department of Biology, University of Padova, Via Ugo Bassi 58 B, 135131 Padova, Italy

  A couple of weeks after the start of my service as President of the International Commission on Zoological Nomenclature, a paper of mine appeared in this Bulletin (Minelli, 1995) in which I outlined some major challenges to be faced by biological nomenclature in response to the changing paradigms of biological systematics. Now, with the approaching end of my six-year term as President, I wish to look in retrospect at the recent history of the Commission, briefly touching on successes and failures alike, and to outline what I now see, from the vantage point of my experience, as further challenges and responsibilities awaiting us; by `us' I mean not merely the Commission but all those active in the field of zoological (or biological) taxonomy.
  The main message I will try to convey in this open letter is that a major effort is necessary in order to better integrate the Commission's work into the daily practice of taxonomists world-wide (and also, in a sense, vice versa).

The Commission and the zoological community
  I have come to realise how poorly known amongst zoologists is the activity (or often even the existence) of the Commission. The Code, despite the thousands of copies printed and sold and the translations now available in several languages, is far from being on the shelves of all practising taxonomists. The Bulletin is only present in a relatively few libraries, mostly in the richer countries, and is largely ignored as a taxonomist's working tool. As a consequence, the rulings which have been issued by the Commission throughout its history are very often overlooked, despite their obvious relevance and the existence of two most useful volumes that provide an index and summary of them (Melville & Smith, 1987; Smith, 2001). Questions of nomenclature are sometimes discussed by zoologists, on a personal level, with ICZN members, but this usually happens because the latter are individually known and appreciated within the local, national or taxonomically specialist community as being knowledgeable in these matters, rather than because of their official affiliation with the Commission. Many more queries are addressed to the Commission's Secretariat in London (more numerous indeed than the limited human and material resources available there can readily cope with). Nevertheless, all these questions are just the tip of a huge iceberg of problems of nomenclature floating through the zoological community, sometimes over years and decades without ever being adequately resolved.
  Some steps have been taken, however, in order to increase the public awareness of the importance of zoological nomenclature, thus improving the visibility of the Code and the Commission's activity. It is not irrelevant, in my view, that a journal such as Science devoted to the fourth edition of the Code a two-column article in its 7 January 2000 issue (Pennisi, 2000), that is in the very week the new Code came into effect. Neither is it irrelevant that an appreciated and widely read monthly such as Trends in Ecology and Evolution hosted my paper on The names of animals in its December 1999 issue (Minelli, 1999). The organizers of the XVIII International Congress of Zoology (Athens, August 2000) selected nomenclature as subject for a General Discussion session that was more than satisfactorily attended. The titles and Abstracts of applications and Commission rulings published in the Bulletin are now displayed on the Commission's Website (www.iczn.org) and so are exposed to a very wide audience.
  The time is ripe for `renegotiating' the relationships between the Commission and the zoological community. I am not speaking of constitutional matters, such as ICZN's affiliation with the International Union of Biological Sciences - in this area, I do not see any reason for proposing changes to the current state of affairs - but of the question: how should the problems of nomenclature be addressed in the near future? To better explain my mind, let me deal with new names and old names separately.
  Up to now, the Commission has only dealt with new names when writing or updating the relevant provisions in the Code; those rules are then placed in taxonomists' hands and implementation becomes the responsibility of individual workers. Each year thousands of new names thus enter zoological nomenclature through the most diverse bibliographic outlets, some of them exceedingly obscure. A feedback from this activity of taxonomists world-wide will reach the Commission only if, and when, names are found to involve problems which individuals cannot (or do not know how to) solve for themselves. In the meantime, of course, those names have become, in their turn, old names.
  Two sets of questions then arise. First, are there any means to improve the way problems with old names are currently addressed by the Commission? Would it be possible to actively involve the whole zoological community in this process? Second, should the Commission take any active role in the `production' of new names? Specifically, should (or could) the Commission be involved in any future system of name registration? My answer to all these questions is yes. I will briefly try to explain why I think so.

Availability of tools
  The Code and the rulings issued by the Commission in response to submitted problems are, or should be, basic tools for all work in zoological taxonomy. For instance, the latest Code is notable in that it allows individuals to take actions to maintain the prevailing usage of names in many circumstances which previously would have needed formal decisions by the Commission.
  With the publication of the fourth edition of the Code, some steps have been taken in order to increase its public accessibility. The production of texts in various languages has been strongly encouraged and, for the first time, all of them are equal in authority. At the time of writing the Code is available in English, French, German, Japanese, Russian and Spanish, and Chinese and Ukrainian texts are in an advanced state of preparation; others may follow. There are proposals to produce the Code on CD, supported by adequate searching software, and for a companion or guide which will be easier to understand than the complex wording of the existing Code.
  Beyond this, however, I should like to mention two more advanced targets.
  The first is a future availability of the Code to any user, free of all cost. The only impediment to the adoption of this policy has been, and still is, the precarious financial position of the Commission. The revenue from the sales of the Code and subscriptions to the Bulletin are, at present, the main source of income which permits the existence of its publications and Secretariat. Technically, it would be easy to have an electronic version of the Code, with searching software, placed on a freely accessible Website. Changes to the financial basis must be made as soon as possible, and indeed might actually happen if the scientific community becomes really aware of the service being offered to it by the Commission and its Secretariat.
  My second belief is that in the near future we should try to re-write the Code itself in a much simpler and more user-friendly way than has been traditional. I am not speaking here of changes in what makes a name available or valid, or in the application of the principle of priority (or other principles) to homonyms or synonyms; nor of the role and power of the Commission to deal with the relatively rare cases which are controversial or where the Code does not provide a solution. I am speaking instead of the way the rules have been presented in the successive Code editions. The obvious desirability of producing a guide to the Code, or of developing simple and powerful searching tools by which to electronically find the Articles of relevance to a particular problem, means in my view that the current rules could be written in a more straightforward way. The price to pay, of course, would be a break with the traditional layout, but this should not imply a break with established rules of nomenclature. Of course, these rules can be changed, and no doubt some will be, but that is another subject.
  Together with the free availability of the Code, I hope we shall be able to offer the zoological community free and easy access to all the Opinions and other rulings issued by the Commission in its 106 years of existence. Digitalization of all relevant documents is currently being considered.

Discussion of cases
  At its meeting held in Athens in August 2000 the Commission discussed at length (see BZN 57: 202-206) procedures which might be adopted in the near future, in order to render the discussion of cases more effective, that is (i) with a larger involvement of the zoological community and (ii) with some degree of decentralization in the management of the discussion, thus obtaining (iii) a substantial reduction in the average time between the submission of the case and the Commission's decision. A small committee has continued working on this subject and the results of this exploration, jointly with the Trust (not-for-profit company) set up in the U.K. to administer the Commission's financial affairs, will be available shortly. With my Presidential term expiring, I must refrain from promising the adoption of any specific measure. Nevertheless, it is probable that the Commission will quickly move towards a large use of internet facilities for the discussion of cases. Specialist nomenclature committees of international zoological societies and qualified internet discussion groups may be co-opted by the ICZN for handling individual cases, in order to provide a richer and better argued documentation to be forwarded to the Commission for its eventual ruling. Involving more people in the actual discussion of cases may open the way to a larger involvement of the zoological community also in other vital events in the Commission's life, such as the election of new Commissioners.

Registration of new names

  I should like to come back to the subject of names for newly recognised taxa. The Code Discussion Draft which was widely circulated in 1995 included proposed rules for the registration of all new names. The public discussion elicited by that document demonstrated that the concept of registration was generally acknowledged as valuable, even though some were opposed in principle. However, the further work of the Commission and its Editorial Committee, in the light of the comments provided by many zoologists from different countries, proved that it was not possible to retain those rules in the final text of the fourth edition of the Code because of difficulties in finding an acceptable way to implement registration. This is not the place to examine the different options we discussed, or the objections raised to them. What truly matters is to stress that the Commission has never abandoned the hope of making registration into a fact, so that all names become visible rather than being buried in the steadily growing mountain of mostly inaccessible publications. I am firmly convinced of the need for the Commission to be closely involved with any form or mechanism of registration, whatever kind of agency will be eventually responsible for it. Very interesting suggestions for cooperation in this respect have been advanced by the publishers of Zoological Record, an acknowledged primary tool of every active taxonomist.

New taxonomies and nomenclatures
  The Commission cannot (and does not!) quietly sleep in an ivory tower without paying attention to deveopments around it, whether actual or suggested, and in the recent literature much has been written of `challenges' to the traditional zoological Code and, equally, to its botanical companion.
  When I became ICZN President there was much interest and apprehension about the `BioCode' initiative (see BZN 53: 148-166, September 1996). Several drafts of a unified Code to be eventually adopted for new names of all organisms (bacterial, botanical and zoological) were produced and some lively bursts of discussion developed at international meetings and on the internet. The project was eventually abandoned, mainly owing to manifest difficulties in satisfactorily dealing with already existing names and to unwillingness of many botanists and zoologists alike to part with their traditional rules and to accept registration of new names. For all those who took part in writing or discussing the BioCode drafts, however, this was a useful experience in that it invited a closer study of the long experience underlying the other Codes, and it brought about some minor but useful convergences in the most recent editions of these and to the establishment of a standing International Committee on Bionomenclature to facilitate liaison between the bodies responsible for the Codes (see BZN 58: 6-7).
  Things are very different with the `PhyloCode' (see Forey, 2001), not just because this initiative is still being actively pursued by its proponents and is the subject of sustained debate, but because of the basic conceptual issues involved in the contrast between the traditional (`Linnaean') Codes and the proposed PhyloCode. In my 1995 paper, I wrote that `We must expect that the development of cladistics will increasingly ask for a revised biological nomenclature', and this is exactly what is happening with the PhyloCode. To be sure, to be a cladist does not automatically mean to be in favour of the new proposal. For example, this has been strongly criticized by cladists such as Nixon & Carpenter (2000) and Forey (2001), who do not subscribe to this plea for abandoning Linnaean nomenclature. The next few years will be interesting in this respect.
  I like the challenges and the debates, but I cannot conclude other than by repeating the closing words of my 1999 paper, that is: `One can imagine that in the future Linnaean and not-Linnaean classification may exist side-by-side. Or maybe not. At any rate, the publication of the new zoological Code could be a good opportunity to open the debate. Otherwise, both parties are likely to go astray: Linnaean-style taxonomists on one side, patiently continuing to produce names that others may be unwilling to use, and phylogenists on the other, perhaps too ready to change the rules. It took one century from Linnaeus to the Strickland Code, and another sixty years to the Regles. Let's talk to one another. Rules can still evolve but a Code, historically, follows and consolidates practice. It does not establish it from scratch.'
  But the dialogue we need to develop is not just the dialogue between the `phylocoders' and the defenders of Linnaean nomenclature. Starting from a common awareness of the importance of biological systematics, of which nomenclature is a humble but necessary arm, we must all cooperate in developing a common strategy in order to raise, internationally as well as locally, the institutional and financial support that systematic biology fully deserves (Boero, 2001). With a better supported taxonomy, with ICZN's activities much more closely intertwined with taxonomic research than they are at present, and with a much better use of internet facilities, our old Commission should be able to adequately fulfil, well into the new century, its institutional role at the service of zoology.

A personal note
  Let me close on a more personal note.
  During my six-year term as President there has been a substantial turnover in the Commission's membership. Of the 27 members present at the beginning, 15 (F.M. Bayer, L.R.M. Cocks, J.O. Corliss, G. Hahn, O. Halvorsen, D. Heppell, L.B. Holthuis, Z. Kabata, P.T. Lehtinen, LW.B. Nye, J.M. Savage, R. Schuster, Y.I. Starobogatov, V.A. Trjapitzin and S.-I. Ueno) retired or left between then and 2000. I wish to thank all of them once more for their valuable contribution to the Commission's work.
  In May this year David Ride retired from the Commission after 38 years of service. Twice President, and editorial Chairman of the third (1985) edition of the Code, David chaired very sensibly and competently the Editorial Committee for the fourth edition. We worked in very close contact between 1996 and 1999, until the new Code was eventually released to the printer. To David I wish to renew the most sincere thanks and appreciation, of the whole Commission and personally mine, for his unique commitment to the Commission and the Code.
  I wish also to extend my words of thanks and appreciation to four more people who will retire soon from their very long and productive association with the Commission: in alphabetic order, Harold Cogger, ICZN Vice-President for many years, including the first ones of my term; Claude Dupuis, most perceptive and careful textual critic and wordsmith; Otto Kraus, my energetic predecessor as President; and Philip Tubbs, our knowledgeable and enthusiastic Secretary for 16 years. All were members of the Code Editorial Committee.
  I should also like to thank Jeremy Smith and Anthea Gentry, members of the Secretariat, for their long, loyal and invaluable service to the work of the Commission.
  The Commission's membership has not just registered losses. Seventeen new members (M.A. Alonso-Zarazaga, W. Bohme, D.J. Brothers, DR. Calder, W.N. Eschmeyer, N.L. Evenhuis, R.A. Fortey, R.B. Halliday, LM. Kerzhner, G. Lamas, S.F. Mawatari, P.K.L. Ng, L. Papp, D.J. Patterson, G. Rosenberg, D.X. Song and J. van Tol) have been elected between 1996 and now, representing fourteen different countries, all continents and a wide range of taxonomic fields. Some of them have already had the opportunity of contributing to the Commission's work in a very substantial way. With them, and with the remaining members of the `old guard', is the challenge of placing zoological nomenclature on an increasingly sounder footing, with the active involvement of the whole zoological community.

Acknowledgements
  I am grateful to Bill Eschmeyer and Philip Tubbs for sharing with me their views, not necessarily coinciding with my own, on most of the items mentioned in this article.

References
Boero, F.
2001. Light after dark: the partnership for enhancing expertise in taxonomy. Trends in Ecology and Evolution, 16: 266.
Nixon, K.C. & Carpenter, J.M. 2000. On the other `phylogenetic systematics'. Cladistics, 16: 298-318.
Forey, P.L. 2001. The PhyloCode: description and commentary. Bulletin of Zoological Nomenclature, 58: 81-96.
Melville, R.V. & Smith, J.D.D. (Eds.). 1987. Official Lists and Indexes of names and works in zoology. 360 pp. International Trust for Zoological Nomenclature, London.
Minelli, A. 1995. The changing paradigm of biological systematics: new challenges to the principles and practice of biological nomenclature. Bulletin of Zoological Nomenclature, 52: 303-309.
Minelli, A. 1999. The names of animals. Trends in Ecology and Evolution, 14: 462-463.
Pennisi, E. 2000. Zoology naming rules eased. Science, 287: 26.
Smith, J.D.D. (Ed.). 2001. Official Lists and Indexes of names and works in zoology. Supplement 1986-2000. 136 pp. International Trust for Zoological Nomenclature, London.

 

General Article

Corrections of authorship and date for gastropod (Mollusca) family-group names placed on the Official List and Official Index

Philippe Bouchet and Jean-Pierre Rocroi
Museum national d'Histoire naturelle, SS rue Buffon, 75005 Paris, France (e-mail: bouchet@mnhn.fr)

Abstract. Of the 90 gastropod (Mollusca) family-group names placed on the Official List (76 names) or the Official Index (14 names), 36 have been entered with an erroneous source of original publication, resulting in the name being listed with wrong author and/or date. The present paper rectifies errors in the List (29 names) and the Index (7 names). The corrections change the date of precedence of the 29 erroneously cited potentially valid names on the List by 1 to 54 years, but, as far as we have ascertained, this will not cause nomenclatural instability. We determined that the remaining 54 gastropod family-group names on the Official List or Index were listed with current references. 

  There are currently 90 gastropod family-group names placed on the Official List or the Official Index as recorded in the volume Official Lists and Indexes of Names and Works in Zoology published in 1987 and the Supplement 1986-2000 published in 2001. In the course of compiling a nomenclator of molluscan supraspecific names, we have discovered that a high proportion (40%) of these names have been entered with an erroneous source of original publication, resulting in the name being listed with wrong author and/or date.
  Under the third (1985) edition of the Code, `a name entered in an Official List [was] deemed to have any authorship, date, publication, name-bearing type, and additional qualification (such as precedence) attributed to it in the relevant Opinion or Direction' (Article 78f(iii)). A correction to the List or Index could not be made as routine book-keeping work, but necessitated a vote by the Commission under the plenary power. To conform with the provisions of the Code, the present list of corrections was submitted as an application in October 1997 (Case 3056, receipt announced in BZN 55: 1), but has not been published. In the meantime, the fourth edition of the Code came into force on 1 January 2000; it states that `the status of a name entered in an Official List is subject to the ruling(s) in any relevant Opinion(s)' (Article 80.6), but also that `Official corrections to errors and omissions ... may be published by the Commission without further vote . . .' (Article 80.4).
  The purpose of the present article is to rectify the errors in the List (29 names) and the Index (7 names), and the Bulletin of Zoological Nomenclature is the most appropriate forum to publish these corrections in a consolidated manner. In addition, we also consider one of the names (CUTHONIDAE) placed on the List in 1966 (Opinion 773) for which the criteria of availability at the time of its original publication had not been met. We determined that the remaining 54 gastropod family-group names in the Official List or Index were listed with proper references. The corrections change the date of precedence of the 29 erroneously cited potentially valid names on the List by 1 to 54 years. However, as far as we have ascertained, the corrections will not cause nomenclatural instability.
  Names on the Official List are in bold type and those on the Official Index are in non-bold type.

ACANTHINULINAE
LIST: (Direction 27, 1955) Pilsbry, 1926, in Tryon, Manual of Conchology. (2)27: 186.
CORRECTION: Steenberg, 1917 [5 October], Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 69: 14.

ACICULIDAE
LIST: (Opinion 344, 1955) Woodward, 1854, Manual of the Mollusca. London, Weale: 178.
CORRECTION: Gray, 1850, Figures of Molluscous Animals, 4: 121. (Original spelling ACICULADAE).

ACMAEIDAE
LIST: (Opinion 344, 1955) Carpenter, 1857, Catalogue of the Reigen Collection of Mazatlan Shells in the British Museum: 202.
CORRECTION: Forbes, 1850, Report of the 19th Meeting of the British Association for the Advancement of Science (Birmingham, 1849), Notices and Abstracts of Communications: 76 (Original spelling AcMAEADAE).

AEOLIDIIDAE
LIST: (Opinion 779, 1966) d'Orbigny, 1834 [sic, should be 1839], Mollusques Echinodermes, Foraminiferes et Polypiers, recueillis aux Iles Canaries. In Webb & Berthelot, Histoire Naturelle des Iles Canaries, Zoologie: 42.
CORRECTION: Gray, 1827, Plates to Zoology: plate Mollusca. Vol. 7, pl. 3 in: Smedley, Rose & Rose (Eds.), Encyclopaedia Metropolitana. (Original spelling EOLIDAE)

AKERIDAE
LIST: (Opinion 539, 1959) Pilsbry, 1893, in Tryon, Manual of Conchology, 15: 350.
CORRECTION: Mazzarelli, 1891, Zoologischer Anzeiger, 14: 243. (Original spelling Aceridae, based on Acera, an incorrect subsequent spelling (Opinion 539) of Akera O.F. Muller, 1776).
REMARKS: It should be noted that Pilsbry (1893, p. 350) himself wrote `Subfamily AKERIDAE (ACERIDAE Mazzarelli)', although, strangely enough, he considered himself to be the author of AKERIDAE.
Early nineteenth century authors used the vernacular family name `Aceres'. Latreille (1825, p. 177) first latinized it as `ACERA', but it did not include, explicitly or implicitly, Akera. The latinized family name `ACERA (Aceres, Cuv.)' was used by Menke (1830, p. 12) and explicitly included `Akera, Cuv.'. However, Cuvier (1810, p. 1) merely used the vernacular `Aceres' to designate gastropods without tentacles. This was a descriptive term opposed to `Diceres' (gastropods with two tentacles) and `Tetraceres' (gastropods with four tentacles), first proposed by Blainville (1816, p. 52), and later latinized (Blainville, 1825, p. 487), but not based on an available generic name. Usage of the descriptive terms `Aceres', `Diceres' and `Tetraceres', or of their latinized form, was abandoned after 1840, but the first valid introduction of a family-group name based on Akera was by Mazzarelli (1841). No current usage attributes AKERIDAE to any of these early 19tb century authors.

AMPHIBOLIDAE
LIST: (Opinion 479, 1957) H. & A. Adams, 1855, The Genera of Recent Mollusca, 2: 268.
CORRECTION: Gray, 1840 [16 October], Synopsis of the Contents of the British Museum, (Ed. 42): 128, 149.

APLYSIIDAE
LIST: (Opinion 1182, 1981) Swainson, 1840, A Treatise on Malacology: 247, 248, 251.
CORRECTION: Lamarck, 1809, Philosophic Zoolagique, 1: 320 [as the vernacular `les Laplysiens', latinized as LAPLYSIANA by Children, 1823, Quarterly Journal of Science, Literature & the Arts, 15: 232]. Also introduced [as LAPLYSIANA] by Rafinesque, 1815, Analyse de la Nature: 142,
Current usage. APLYSIDAE is attributed to Lamarck, 1809 by, e.g., Franc (1968, p. 849), Nordsieck (1972, p. 42), Ros (1975, p. 307), Cattaneo & Barletta (1984, p. 203), Cervera et al. (1988, p. 19), Ortea & Martinez (1990, p. 17), and Sabelli et al. (1992, p. 436); it is attributed to Swainson, 1840, with reference to Opinion 1182 by Hoisaeter (1986, p. 103); and to Rafinesque, 1815 by, e.g., Abbott (1974, p. 342), Arakawa & Hoshino (1982, p. 134), Rios (1985, p. 181), Fukuda (1992, p. 75), and Tracey et al. (1993, p. 155). There is thus no single `generally accepted' (in the sense of Article 11.7.2) usage of the author and date of APLYSIIDAE. Attributing it to Lamarck appears to be the parsimonious option.

AZECINAE
LIST: (Direction 27, 1955) Kennard & Woodward, 1926, Synonymy of the British non-marine Mollusca: xvi, 144.
CORRECTION: Watson, 1920 [2 May], Proceedings of the Malacologieal Society of London, 14(1): 24.

CHROMODORIDIDAE
LIST: (Opinion 1375, 1986) Bergh, 1892, Malacologische Untersuchungen in: Reisen im Archipel der Philippinen von Dr. C Semper, Theil, Wissenschaftliche Resultate. Section 2, vol. 3, part 18, p. 1103.
CORRECTION: Bergh, 1891 [October], Zoologische Jahrbucher (Abteilung fiir Systematik, Geographie und Biologic der Thiere), 6: 137.

COMINELLINAE
LIST: (Opinion 479, 1957) P. Fischer, 1884, Manuel de Conchyliologie, 7: 624.
CORRECTION: Gray, 1857 [9 May], Guide to the systematic distribution of Mollusca in the British Museum. Part I: 15.

CUTHONIDAE
LIST: (Opinion 773, 1966) Odhner, 1934, British Antarctic (Terra Nova) Expedition 1910, Natural History Report, Zoology. 7(5): 278.
REMARKS: The name CUTHONIDAE was introduced without a description and was not available under Article 13a of the Code at the time of Opinion 773 [now Article 13.1]. It was first diagnosed by Odhner, 1939, Det Kongelige Norske Videnskabers Selskabs, Skrifter, 1939(1): 53. The name CUTHONIDAE satisfies the conditions of Article 13.2.1 and is available from Odhner (1934).

HELICARIONIDAE
LIST: (Opinion 1678, 1992) Bourguignat, 1883, Annales des Sciences Naturelles, (Zoologie). Art. 2, (6)15: 9.
CORRECTION: Bourguignat, 1877, Bulletin de la Societe des Sciences Physiques et Naturelles de Toulouse, 3: 64. A correction to Godwin-Austen, 1882, Land and freshwater Mollusca of India, Part 2: 65 was notified in BZN, 50(4): 313 and published in the 2001 Supplement to the Official Lists and Indexes.

HELICELLINAE [based on Helicella Ferussac, 1821]
LIST: (Opinion 431, 1956) Hesse, 1926, Archiv fur Molluskenkunde, 58(3):115. A correction to Ihering, 1909, Verhandlungen der zoologisch-botanischen Gesellschaft in Wien, 1909: 429 was published in the 1987 Official Lists and Indexes.

HELICELLINAE (based on Helicella Lamarck, 1812, not an available name) INDEX: (Opinion 431, 1956) Chenu, 1859, Manuel de Conchyliologie, 1: 421. CORRECTION: H. & A. Adams, 1855 [January], The Genera of Recent Mollusca, 2: 112.

HYGROMIINAE
LIST: (Direction 27, 1955) Geyer, 1909, Unsere Land- and Susswasser-Mollusken. (Ed. 2): 11.
CORRECTION: Tryon, 1866 [6 October], American Journal of Conchology, 2(4): 306.

LAURIINAE
LIST: (Direction 27, 1955) Thiele, 1931, Handbuch der systematischen Weichtierkunde, 1(2): 509.
CORRECTION: Steenberg, 1925 [18 June], Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 80: 201.

LYMNAEIDAE
LIST: (Opinion 495, 1957) Rafinesque, 1815, Analyse de la Nature: 144.
COMMENT: Lamarck, 1812, Extrait du cours de zoologie: 116 established the vernacular name `les Lymneens' which was subsequently latinized (as LYMNAEANA) by Children, y 1823, Quarterly Journal of Science, Literature & Arts, 15: 242. Although the name LYMNAEIDAE has generally been credited to Lamarck in the 19th century literature, current usage refers this family name to Rafinesque (e.g., Franc, 1968, p. 528; Starobogatov, 1970, p. 46; Tracey et al., 1993, p. 158; Giusti et al., 1995, p. 169). It may appear inconsistent to accept some of Lamarck's vernacular names (see, e.g., APLYSIIDAE and TRITONIDAE herein, and many bivalve family-group names in current use) and reject others. However, names are to be evaluated individually on a case by case approach and on these grounds it seems justified to accept the author and date of LYMNAEIDAE

NARICIDAE
INDEX: (Opinion 1009, 1974) Recluz, 1846, [`1845'], Magasin de Zoologie, (2)7(9): 6. CORRECTION: The author and reference are correct, but the citation of the date suggests that the work was published in 1846 rather than in the nominal year, 1845. This does not appear to be correct. The folios containing Recluz' work are dated `Octobre 1845' on the last printed line of the first page starting the folio, and a copy in the mollusc library of the Museum national d'Histoire naturelle in Paris bears a manuscript note in the upper left corner of page 49: `1er` Xbre' [=1 October]. The Secretariat does not hold any information that justifies the date 1846 and the date 1845 should be accepted. NARICIDAE is a junior objective synonym of VANIKORIDAE Gray, 1840.

OTINIDAE
LIST: (Direction 27, 1955) Chenu, 1859, Manuel de Conchyliologie, 1: 479.
CORRECTION: H. & A. Adams, 1855 [September], The Genera of Recent Mollusca, 2: 249.

PALUDINIDAE
INDEX: (Opinion 573, 1959) Gray, 1840, Synopsis of the contents of the British Museum. Ed. 42: 117.
CORRECTION: Fitzinger, 1833, Beitrkge zur Landeskunde Oesterreich's unter der Enns, 3: 109 (as `Gruppe' Paludinoidea). First used as the vernacular `Les Paludinides' by Risso, 1826, Histoire Naturelle ... de 1'Europe Meridionale, 4: 100, but there is no current usage to attribute the name to Risso. PALUDINIDAE is based on the name Paludina Ferussac, 1812 (a junior objective synonym of Viviparus Montfort, 1810).

PELTIDAE
INDEX: (Opinion 811, 1967) Winckworth, 1931, Plymouth Marine Fauna. (Ed. 2): 267.
CORRECTION: Vayssiere, 1885, Annales du Musee d'Histoire naturelle de Marseille, Zoologie, 2(3): 104.

PLANORBIDAE
LIST: (Direction 27, 1955) Gray, 1840, in Turton, Manual of the land and fresh-water shells of the British Islands. (Ed. 2): 256.
CORRECTION: Rafinesque, 1815, Analyse de la Nature: 143. Original spelling (subfamily) PLANOxBIA (of the family TROCHINIA).
REMARKS: Rafinesque based the family-group name PLANORBIDAE on `Planorbis Geof.'. This is Planorbis of Geoffroy (1767, p. 12), a work placed on the Official Index of Rejected and Invalid Works by Opinion 362. Planorbis was first made available by O.F. Muller (1774, p. 152), who refers explicitly (e.g., pp. 154, 157, 159, 160, etc.) to Geoffroy, so that the reference by Rafinesque to `Planorbis Geof.' unambiguously denotes the taxon now attributed to Muller. The family PLANORBIDAE is attributed to Rafinesque, 1815 in many current publications, following H.B. Baker (1956, p. 133). By attributing PLANORBIDAE to Rafinesque, 1815, this name takes the same precedence as the name ANCYLIDAE Rafinesque, 1815 (placed on the Official List by Direction 41). Starobogatov (1967, p. 293) acted as First Reviser and gave relative precedence to the name PLANORBIDAE over ANCYLIDAE Rafinesque, 1815, which he attributed to the subfamily ANCYLINAE within the PLANORBIDAE.

PURPURIDAE
LIST: (Opinion 886, 1969) Broderip, 1839, Penny Cyclopaedia, 14: 321.
CORRECTION: Children, 1823, Quarterly Journal of Science, Literature & the Arts, 16: 54 [original spelling PURPURIFERA, latinization of the vernacular `les Purpuracees' of Lamarck, 1809, Philosophie Zoologique, 1: 322]. Current usage. In Opinion 886 THAIDIDAE/INAE Jousseaume, 1888 was given precedence over PuRPUxIDAE/INAE (there attributed to Broderip, 1839). Attributing the name PURPURIDAE to Lamarck (1809) would cause nomenclatural instability because PURPURIDAE would then have precedence over MURICIDAE Rafinesque, 1815. THAIDINAE is used for a subfamily within MuRICIDAE Rafinesque, 1815.

REALIINAE
INDEX: (Opinion 973, 1971) Pfeiffer, 1858, Monographia Pneumonopomorum Viventium, suppl. 1: 153.
CORRECTION: Pfeiffer, 1853 [12 February], Catalogue of Phaneropneumona or Terrestrial Operculated Mollusca in the Collection of the British Museum: 217 [original spelling REALIANA].

RETUSIDAE
LIST: (Opinion 568, 1959) Thiele, 1931, Handbuch der systematischen Weichtierkunde, 1: 189 [sic, error for p. 389].
CORRECTION: Thiele, 1925 [before 10 November], Deutsche Tiefsee-Expedition 1898-1899, 17(2): 234.

RUMINIDAE
LIST: (Direction 27, 1955) Thiele, 1931, Handbuch der systematischen Weichtierkunde, 1(2): 554.
CORRECTION: Wenz, 1923 [5 June], Fossilium Catalogus, I, Pars 20: 875.

RUNCINIDAE
LIST: (Opinion 811, 1967) Gray, 1857, Guide to the systematic distribution of Mollusca in the British Museum. Part I: 204.
CORRECTION: H. & A. Adams, 1854 [October], The Genera of Recent Mollusca, 2: 42.

SCHIZOSTOMIDAE
INDEX: (Opinion 1470, 1988) Eichwald, 1871, Geognostica paldeontologische Bemerkungen, uber die Halbinsel Mangischlak und die Aleutischen Inseln: 119.
CORRECTION: Bronn, 1849, Index Palaeontologicus, 2(B): 421 [original spelling SCHIZOSTOMICA].

STRUTHIOLARIINAE
LIST: (Opinion 479, 1957) P. Fischer, 1884, Manuel de Conchyliologie: 677. CORRECTION: Gabb, 1868 [3 November], American Journal of Conchology, 4(3): 147 [original spelling STRUTHIOLARINAE].

SUBULINIDAE
LIST: (Direction 27, 1955) Thiele, 1931, Handbuch der systematischen Weichtierkunde, 1(2): 549.
CORRECTION: Fischer & Crosse, 1877, Mission scientifique au Mexique et dans Z'Amerique Centrale. Recherches Zoologiques (7), 1(6): 592.

THAIDIDAE
LIST: (Opinion 886, 1969) Suter, 1913, Manual of the New Zealand Mollusca: 420. A correction to Jousseaume, 1888, Memoires de la Societe Zoologique de France, 1: 179 [original spelling THAISIDAE] was published in the 1987 Official Lists and Indexes.

TRITONIIDAE
LIST: (Opinion 668, 1963) H. & A. Adams, 1858, The Genera of Recent Mollusca, 2: 62.
CORRECTION: Lamarck, 1809, Philosophie Zoologique, 1: 320 (as the vernacular `les tritoniens', latinized as TRITONIANA by Children, 1823, Quarterly Journal of Science, Literature & the Arts, 15: 222).
Current usage. TRITONODAE is attributed to Lamarck, 1809 by, e.g., Franc (1968, p. 873), Nordsieck (1972, p. 65), Ros (1975, p. 332), Schmekel & Portmann (1982, p. 141), McDonald (1983, p. 120), Hoisaeter (1986, p. 107), Bertsch & Mozqueira (1986, p. 47), Cervera et al. (1988, p. 38) and Rolan et al. (1991, p. 113); it is attributed to H. & A. Adams, 1858 by, e.g., Sabelli et al. (1992, p. 446) and Cattaneo-Vietti et al. (1990, p. 22); and to Menke, 1828 by, e.g., Abbott (1974, p. 368) and Rios (1985, p. 188). There is thus no single `generally accepted' (in the sense of Art. 11.7.2) usage of the author and date of TRITONIIDaE. Attributing it to Lamarck appears to be the parsimonious option.

TURBINELLIDAE
LIST: (Opinion 489, 1957) Swainson, 1840, A Treatise on Malacology: 75. CORRECTION: Swainson, 1835, The elements of modern conchology: 13, 20.

VALLONIIDAE
LIST: (Direction 27, 1955) Pilsbry, 1900, Proceedings of the Academy of Natural Sciences of Philadelphia, (1900): 564.
CORRECTION: Morse, 1864 [17 March], Journal of the Portland Society of Natural History, 1: 5, 21 [original spelling (subfamily) VALLONINAE].

VERTIGINIDAE
LIST: (Direction 27, 1955) Stimpson, 1851, Shells of New England: 53.
CORRECTION: Fitzinger, 1833, Beitrkge zur Landeskunde Oesterreich's unter der Enns, 3: 109 (original spelling (`Gruppe') VERTIGINOIDEA).

VITREINAE
LIST: (Direction 27, 1955) Thiele, 1931, Handbuch der systematischen Weichtierkunde, 1(2): 587.
CORRECTION: H.B. Baker, 1930 [24 April], The Nautilus, 43(4): 122.

XANCIDAE
INDEX: (Opinion 489, 1957) Woodring, 1928, Carnegie Institution of Washington, Publication No. 385: 250.
CORRECTION: Pilsbry, 1922 [4 January], Proceedings of the Academy of Natural Sciences of Philadelphia, 73: 342.

XENOPHORIDAE
LIST: (Opinion 715, 1964) Philippi, 1853, Handbuch der Conchyliologie und Malacozoologie: 185.
CORRECTION: Troschel, 1852, Archiv fur Naturgeschichte, 18(2): 280 [original spelling XENOPHORACEA].

Acknowledgements
  We are grateful to a number of colleagues who have drawn our attention to little-known sources of molluscan family-group names and who have provided photocopies of rare references, notably R. Burn (Geelong, Australia), G. Falkner (Worth-Horlkofen, Germany), A. Kabat (Washington, D.C., U.S.A.), J. Pickering (London, U.K.). A. Kabat and G. Rosenberg (Philadelphia, U.S.A.) read the manuscript before publication and agreed with the conclusions.

References
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